Open access papers

Below are links to pdf files of published papers under the respective categories

Taxonomy and nomenclature
New records and new reports
Phylogenetics and evolution

Taxonomy and Nomenclature

D. Schill & D. Long. 2002. Anastrophyllum lignicola (Lophoziaceae), a new species from the Sino-Himalaya, and A. hellerianum new to China. Ann. Bot. Fennici 39: 129-132

A.D. Potemkin. 2001. Three new species of Scapania (Hepaticae) from India and China. Ann. Bot. Fennici 38:83-89

X-L. He. & R. Grolle. 2001. Xylolejeunea, a new genus of the Lejeuneaceae (Hepaticae) from the Neotropics, Madagascar and the Seychelles. Ann. Bot. Fennici 38:25-44

R. Grolle & R.-L. Zhu. 1999. Drepanolejeunea longii (Lejeuneaceae, Hepaticae), a new species from Bhutan. Ann. Bot. Fennici 36:115-118

A.D. Potemkin. 2000. Bryophyte flora of Hunan Province, China. 2. Scapania koponenii sp. Nova (Scapaniaceae, Hepaticae) Ann. Bot. Fennici 37:41-44

Zhu, R.-L. & So, M. L. 1998, Notes on the taxonomy and distribution of Cololejeunea platyneura (Hepaticae, Lejeuneaceae), Annales Botanici Fennici 35: 229-232


Gökler, I. & Aysel, V. 1998. A New Aquatic Liverwort for the Flora of Turkey. 
Rycroft, D.S., Cole, J. W., Heinrichs, J., Groth, H., Renker, C. Pröschold, T. 2002. Phytochemical, Morphological, and Molecular Evidence for the Occurrence of the Neotropical Liverwort Plagiochila stricta in the Canary Islands, New to Macaronesia.
Özenoglu, H. & Gökler, I. 2001 [2002], Liverworts (Marchantiopsida) of the Dilek Peninsula National Park, Turkish Journal of Botany 26: 297-301. 
Çetin, B. 1998, The liverworts (Hepaticae) of Sinop and its environs (Ayancik, Boyabat and Gerze), Turkish Journal of Botany 23(2): 113-116. 1 fig. 
Çetin, B. 1999, The liverworts (Hepaticae) of Uludag National Park (Bursa), Turkish Journal of Botany 23(4): 277-280 1 fig. 
Aleffi, M. - Dipartimento di Botanica ed Ecologia, Universita di Camerino. 2001, Checklist of Italian Liverworts, On the web site of University of Trieste, Dept. of Biology 
Berg, Ch. & Wiehle, W. 1991, Rote Liste der Moose (Bryophyta) Mecklenburg-Vorpommerns, Ernst-Moritz-Arndt-Universität Greifswald web page
Boggan, J., Funk, V. A., Kelloff, C. L., Hoff, M., Cremers, G. & Feuillet, C. P. G.-A. 1997, Checklist of the plants of the Guianas (Guyana, Surinam, French Guiana) 2nd Edition, Smithsonian National Museum of Natural History web site
But, P. P.-H. [Ed.]. 2004, Hong Kong Flora and Vegetation. Checklist of Hong Kong Bryophytes (III. Liverworts). Liverworts, Hong Kong Flora and Vegetation web site
Casas, C. 1998, The Anthocerotae and Hepaticae of Spain and Balearic Islands : a preliminary checklist, Orsis 13: 17-26. Updated
Casas, C. 2002, Llista de les espècies d'Anthocerotae i d'Hepaticae d'Espanya i les Illes Balears, Univeristat Autònoma de Barcelona web site
Cavalcanti Pôrto, K., Rangel Germano, S. & Marques Borges, G. 2005, Avaliação dos Brejos de Altitude de Pernambuco e Paraíba, quanto à Diversidade de Briófitas, para a Conservação, Published online by Centro de Pesquisas Ambientais do Nordeste, Brazil 
Christy, J. A. 2003, Bryophytes of Fort Clatsop National Memorial, Oregon, Evansia 20: 107-113
Crosby, M. R. & Magill, R. E. 2004, Index of Bryophytes 2001-2004
Crosby, M. R. & Magill, R. E. 2005, Index of Bryophytes 2005
Dauphin, G. 1999, Bryophytes of Cocos Island, Costa Rica: diversity, biogeography and ecology, Revista de Biología Tropical 47(3): 309-328
de Castro Molinaro, L. & Pinheiro da Costa, D. 2001, Briófitas do arboreto do Jardim Botânico do Rio de Janeiro, Rodriguésia 81: 107-124
Dirkse, G. M., During, H. J. & Siebel, H. N. 1999, Standaardlijst van de Nederlandse blad-, lever- en hauwmossen, Buxbaumiella 50(2): 68-128
Eckel, P. M. 2004, Preliminary Cryptogamic (Moss, Lichen and Liverwort) Flora of the Canadian and American Gorge at Niagara Falls, Res Botanica, a Missouri Botanical Garden Web site, St. Louis, MO 128 pp. 
Erzberger, P. & Papp, B. 2004, Annotated checklist of Hungarian bryophytes, Studia Botanica Hungarica 35: 91-149
Fernandes Peralta, D. & Yano, O. 2005, Briófitas de mata paludosa, município de Zacarias, noroeste do Estado de São Paulo, Brasil, Acta Botanica Brasilica 19(4): 963-977
Frahm, Jan-Peter & Michael Lüth 2008, The Bryophyte Flora of the Maltese Islands, Archive for bryology: 29 
Frahm, Jan-Peter, Michael Lüth & Huub Van Melick 2008, Kommentierte Artenliste der Moose von Sardinien, Archive for bryology: 31
Furuki, T. & Mizutani, M. 2004, Checklist of Japanese Hepaticae and Anthocerotae 2004, Bryological Research 8(10): 296-316
Gökler, I. 1998, Liverworts (Marchantiopsida) of the Altindere Valley National Park, Turkish Journal of Botany 22: 409-412. 1 fig. 1 tab.
Higashino, P. K., Cuddihy, L. W., Anderson, S. J. & Stone, C. P. 1988, Bryophytes and Vascular Plants of Kipahulu Valley Haleakala National Park, University of Hawaii at Manoa Technical Report 65
Hill, M. & Preston, C. D. 2005, British and Irish bryophytes and their distribution, British Bryological Society web page
Hodgetts, N. 2004, Mosses and liverworts of Uganda (MALOU). Checklist of hepatics, U.K. Natural History Museum web site
Hodgson, E. A. 1944, A Review of the New Zealand Species of the Genus Chiloscyphus. Being the Second of a Series of Papers on New Zealand Hepaticae, Transactions and Proceedings of the Royal Society of New Zealand 73: 27-52
Hutten, M., Woodward, A. & Hutten, K. 2005, Inventory of the Mosses, Liverworts, Hornworts, and Lichens of Olympic National Park, Washington: Species List, U.S. Geological Survey Scientific Investigations Report 2005-5240, 78 pp.
Ingerpuu, N., Kalda, A., Kannukene, L., Krall, H., Leis, M. & Vellak, K. (compilers). 2005, List of Estonian bryophytes. Updated in January 2005, Zooloogia ja Botaanika Instituut web site
Jóhannsson, B. 2003, Checklist of bryophytes in Iceland. Skrá yfir mosa á Íslandi, Fjölrit Náttúrufraedistofnunar 44
Kürschner, H. & Erdag, A. [2003]. 2004, Bryophytes of Turkey: An Annotated Reference List of the Species with Synonyms from the Recent Literature and an Annotated List of Turkish Bryological Literature, Turkish Journal of Botany 29: 95-154
Koponen, T., Enroth, J., Fang, Y.-M., Huttunen, S., Hyvönen, J., Ignatov, M., Juslén, A., Lai, M.-J., Piippo, S., Potemkin, A. D. & Rao, P. 2000, Bryophyte flora of Hunan Province, China. 1. Bryophytes from Mangshan Nature Reserve and Wulingyuan Global Cultural Heritage Area, Annales Botanici Fennici 37: 11-39
Kubinská, A. (Editor) 1992?, Slovakian liverworts, Botanicky ústav SAV web site
Kucera, Jan & Vána, J. 2003, Check- and Red List of bryophytes of the Czech Republic 2003, Preslia 75: 193-222
Lyman, K. J. 1999?, Checklist of Wisconsin Liverworts, Milwaukee Public Museum - Section For Botany - web page
McCarthy, P. M. 2006, Checklist of Australian liverworts and hornworts, Australian Biological Resources Study, Canberra. Version 6 April 2006
<No author> 2005, Census catalogue of the Irish liverworts, National Botanic Gardens of Ireland web page
Piippo, S., He, X.-L. & Koponen, T. 1997, Hepatics from northwestern Sichuan, China, with a checklist of Sichuan hepatics, Annales Botanici Fennici 34: 51-63
Piippo, S., He, X.-L., Juslén, A., Tan, B. C., Murphy, D. H. & Pócs, T. 2002, Hepatic and hornwort flora of Singapore, Annales Botanici Fennici 39: 101-127
Pinheiro da Costa, D. 2002 [2003], Florisitic composition and diversity of Amazonian rainforest bryophytes in the state of Acre, Brazil, Acta Amazonica 33(3)
Reiner-Drehwald, M. E. 1999, Catalogue of the genus Lejeunea Lib. (Hepaticae) of Latin America, Bryophytorum Bibliotheca 54. 101 pp. 2 figures, 1 table, 22x14cm
Salazar Allen, N. & Chung, C. 2003, The bryophytes (hornworts, liverworts and mosses) of the Bahia Honda region. (Veraguas, Panamá), LIQUID JUNGLE LAB web site
Siebel, H. N. & al. 2005, Standaardlijst van de Nederlandse blad-, lever- en hauwmossen, Buxbaumiella 73
Singh, Ajit Pratap, Dinesh Kumar and Virendra Nath 2008, Studies on the Genera Frullania Raddi and Jubula Dum. from Meghalaya (India): Eastern Himalayas, Taiwania, 53(1): 51-84
Stark, L. R. & Whittemore, A. T. 2001, Bryophytes from the northern Mojave desert. I. A list of reported species. Final version, Southwestern Naturalist 45: 226-232
Syrjänen, K. 2002, Sammalten taksonomia ja nimistö, The web site of Finland's environmental administration
Söderström, L. & Hedenäs, L. 1998, Checklista över Sveriges mossor, Myrinia 8(2): 58-90
Söderström, L., E. Urmi & J. Vána. 2002, Distribution of Hepaticae and Anthocerotae in Europe and Macaronesia, Lindbergia 27(1): 3-47
Timme, S. L. & Redfearn, P. L. Jr. 1997, Checklist of the liverworts and hornworts of the interior highlands of North America in Arkansas, Illinois, Missouri and Oklahoma, Evansia 14(3): 89-105
van Tooren, B. & Bruin, K. 2004, Bryophytes of the sand dunes of the West Frisian Islands, Lindbergia 29(1): 51
Weber, W. A. & Wittmann, R. C. 2000, Catalog of the Colorado Flora: a Biodiversity Baseline Liverworts: electronic version 11-Mar-00, University of Colorado Museum. Boulder, Colorado. University Press of Colorado
Werner, J. 2003, Checklist and red list of the bryophytes of Luxembourg, Musée national d'histoire naturelle, Luxembourg web site
Wigginton, M. J. 2001, Checklist of Malawi liverworts, U.K. Natural History Museum web site
Wigginton, M. J. 2002, Checklist and distribution of the liverworts and hornworts of Sub-Saharan Africa, including East African islands, Tropical Bryology Research Reports 3: 1-88
Wigginton, M. J. 2004, Checklist and distribution of the liverworts and hornworts of sub-Saharan Africa, including the East African Islands. 2nd ed., Tropical Bryology Research Reports 5: 1-102
Yano, O. & Aguiar Saraiva Câmara, P. E. 2004, Briófitas de Manaus, Amazonas, Brasil, Acta Amazonica 34(3): 445-457
Zhu, R.-L. 2005, New Checklist of Chinese liverworts, hornworts, and takakiophytes. 2nd version in July 2005, East China Normal University web site

New Records and New Reports

R.-L. Zhu & M.L. So. 1999. New records of Cololejeunea (Lejeuneaceae, Hepaticae) for Chinaand Vietnam.  Bot. Bull. Acad. Sin. 40:165-171
I. Goekler & V. Aysel. 1998. A new aquatic liverwort for the flora of Turkey. Tr. J. of Botany 22: 355-357
T.L. Blockeel, T.M. Ros, M.Sabovljevic M.J. Cano, M.T. Gallego & J. Munoz. 2002. New and interesting bryophyte records for Greece.  Crypt.Bryol. 23:149-155
R.-L. Zhu & M.L. So. 1998. Two epiphyllous liverworts, Cololejeunea dozyana (Dande Lac.) Schiffn. And Cololejeunea sigmoidea Jovet-Ast & Tixier (Hepaticae, Lejeueneaceae), new to Taiwan. Bot. Bull. Acad. Sin. 39:125-130
R.-L. Zhu & M.L. So. 1999. Additions of Lejeuneaceae taxa to the hepatic flora of Yunnan, China.  Ann. Bot. Fennici 36:219-229
Goekler, H. Oezenoglu, F. Kiremit. 2000. A new liverwort for the flora of Turkey. I. Turk. J. Bot. 24:81-83
HARALD KÜRSCHNER. 2003. Nineteen new records to the bryophyte flora of Socotra Island. Additions to the Bryophyte Flora of the Arabian Peninsula and Asocotra 5. Wildenowia 33: 445-458


Phylogenetics and evolution


X. He-Nygren & S. Piippo. Phylogenetic relationships of the generic complex Chiloscyphus-Lophocolea-Heteroscyphus (Geocalycaceae, Hepaticae): Insights from three chloroplat genes and morphology. 2003. Ann Bot. Fennici 40:317-329
A.D. Potemkin. 2002. Phylogenetic system and classification of the family scapaniaceae Mig. Emend. Potemkin (Hepaticae)  Ann. Bot. Fennici 39:309-334
He-Nygre´n, X, Jusle´n, A., Ahonen, I., Glenny, D., and Piippo, S. 2006. Illuminating the evolutionary history of liverworts (Marchantiophyta)—towards a natural classification.




AHONEN, I. 2005. Evolutionary relationships of liverworts with a special focus on the order Porellales and the family Lejeuneaceae.

JUSLÉN, A. 2006. The family Herbertaceae and its novel systematic position within liverworts.